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In Arabidopsis, SIR is encoded by a single, essential gene and is described as a “bottleneck” in the sulfate assimilation pathway ( SIR expression is rapidly upregulated in Arabidopsis and tomato in response to higher sulfite content, and accordingly SIR activity increases ( SIR transcripts are also upregulated after cold stress and methyl viologen treatment in maize ( Synechocystis (Figure 2 and Supplementary Table 2)